Peru State College
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Dr. Joanna Cielocha

Contact Information:

Office: Hoyt 306

Office Phone: (402) 872-2231



Office Hours:

Monday: 2:30-3:30 pm

Tuesday: 9:30-10:30 am

Wednesday: 8:30-9:30 am

or by appointment




14 Fall Course Offerings:

Biol 102 (sections 000C) - General Biology & Lab

Biol 210 (sections 000A & 000B) - Human Anatomy & Lab

Biol 424 (section 000A & 000B) - Biology of Endotherms & Lab


15 Spring Course Offerings:

Biol 102 (section 000A) - Introductory Zoology

Biol 301 (section 000A & 000B) - Microbiology & Lab

Biol 312 (section 000A & 000B) - Human Physiology & Lab



By training, I am a systematic parasitologist. My current research focuses on understanding diversity and interrelationships of a group of cestodes (tapeworms) which exclusively parasitizes elasmobranchs (sharks and rays) as adults. As part of this research, I have participated in field collections of elasmobranch tapeworms in the Gulf of Mexico and Atlantic Ocean off the United States and Belize and the Pacific Ocean off Costa Rica and Vietnam. Many of the specimens used in my research have come from ongoing collections from around the world including specimens collected from sharks and rays off the coasts of Africa, Australia, Borneo, India, Japan, Mexico, Solomon Islands, and Thailand.


Motivated undergraduate students who are interested in understanding and studying the diversity and relationships of parasitic organisms are encouraged to come by my office to discuss potential research projects.



  • Cielocha, J. J., A. Yoneva, M. E. Cantino, S. Daniels & Kirsten Jensen. Accepted. Investigation of sperm ultrastructure of Adelobothrium sp. (Cestoda: Lecanicephalidea). Invertebrate Biology.
  • Cielocha, J. J. & K. Jensen. 2013. Stoibocephalum n. gen. (Cestoda: Lecanicephalidea) from the sharkray, Rhina ancylostoma (Elasmobranchii: Rhinopristiformes), from northern Australia. Zootaxa 3626 (4): 558–568. DOI: 10.11646%2Fzootaxa.3626.4.9.
  • Borucinska, J. D., J. J. Cielocha, & K. Jensen. 2013. The parasite-host interface in the zonetail butterfly ray Gymnura zonura, infected with Hexacanalis folifer (Cestoda: Lecanicephalidea). Journal of Fish Diseases 36: 1–8. DOI:10.1111/j.1365-2761.2012.01408.x.
  • Cielocha, J. J. & K. Jensen. 2011. A revision of Hexacanalis Perrenoud, 1931 (Cestoda: Lecanicephalidea) and description of H. folifer from the zonetail butterfly ray, Gymnura zonura (Bleeker) (Rajiformes: Gymnuridae). Systematic Parasitology 79(1): 1–16. DOI 10.1007/s11230-011-9291-1.
  • Cielocha, J. J., T. J. Cook, & R. E. Clopton. 2011. Host utilization and distribution of nubenocephalid gregarines (Eugregarinorida: Actinocephalidae) parasitizing Argia spp. (Odonata: Zygoptera) in the central United States. Comparative Parasitology 78(1):152–160. DOI: 10.1654/4459.1.
  • Clopton, R. E., T. J. Cook, & J. J. Cielocha. 2010. Nubenocephalus nickoli n. sp. and Nubenocephalus xunantunichensis n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae) parasitizing damselflies (Odonata: Zygoptera) in Belize, Central America. Comparative Parasitology 77(2): 125–136. DOI: 10.1654/4438.1.
  • Hays, J. J., R. E. Clopton, T. J. Cook, & J. L. Cook. 2007. Revision of the genus Nubenocephalus and description of Nubenocephalus secundus n. sp. (Apicomplexa:  Actinocephalidae) parasitizing adults of Argia sedula  (Odonata: Zygoptera: Coenagrionidae) in the primitive Texas Big Thicket, U.S.A. Comparative Parasitology 74(2): 286–293. DOI: 10.1654/4287.1.
  • Clopton, R. E. & J. J. Hays. 2006. Revision of the genus Protomagalhaensia and description of Protomagalhaensia wolfi n. comb. (Apicomplexa: Eugregarinida: Hirmocystidae) and Leidyana haasi n. comb. (Apicomplexa: Eugregarinida: Leidyanidae) parasitizing the Lobster Cockroach, Nauphoeta cinerea (Dictyoptera: Blaberidae). Comparative Parasitology 73(2): 137–156. DOI: 10.1654/4241.1.
  • Hays, J. J., R.E. Clopton, D.L. Cappaert, & D.R. Smitley. 2004. Revision of the genus Stictospora and description of Stictospora villani, n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae) from larvae of the Japanese Beetle, Popillia japonica (Coleoptera: Scarabaeidae), in Michigan. Journal of Parasitology 90(6): 1450–1456. DOI: 10.1645/GE-345R.


Ph.D. in Ecology and Evolutionary Biology, The University of Kansas; May 2013

M.S. in Biological Sciences, Sam Houston State University; August 2008

B.S. in Natural Sciences, Peru State College; May 2006